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Furthur research on the genera Lucanus Scopoli and Eolucanus Kurosawa from China,with the description of two new species1

An updated checklist of Chinese species of the genera Lucanus and Eolucanus1

Notes on some species of the genera Lucanus and Eolucanus3

Ⅰ．Lucanus wuyishanensis Schenk武夷深山3

Ⅱ．Lucanus liuyei Huang ＆ Chen刘氏深山4

Ⅲ．Lucanus swinhoei Parry姬深山5

Ⅳ．Lucanus prossi Zilioli普氏深山5

Ⅴ．Lucanus brivioi Zilioli布氏深山6

Ⅵ．Lucanus nobilis Didier雅深山6

Ⅶ．Lucanus dirki Schenk大理深山6

Ⅷ．Lucanus formosus Didier橙深山7

Ⅸ．Lucanus fujitai Katsura ＆ Giang北越深山7

Ⅹ．Lucanus sp．incert．墨脱未定名深山8

Ⅺ．Lucanus delavayi Fairmaire黄鞘深山8

Ⅻ．Lucanus parryi parryi Boileau帕瑞深山指名亚种9

ⅩⅢ．Lucanus fujianensis Schenk微齿姬深山9

ⅩⅣ．Lucanus fairmairei Planet四川深山10

ⅩⅤ．Lucanus laminifer Waterhouse拉叉深山10

ⅩⅥ．Lucanus angusticornis inclinatus Schenk弯叉深山中国亚种11

ⅩⅦ．Lucanus hermani De Lisle巨叉深山11

ⅩⅧ．Lucanus planeti planeti Planet普叉深山指名亚种11

ⅩⅨ．Lucanus planeti dayaoshanensis Schenk普叉深山广西亚种11

ⅩⅩ．Lucanus dybowski lhasaensis Schenk斑股深山四川亚种12

ⅩⅪ．Lucanus ludivinae Boucher路氏深山13

ⅩⅫ．Lucanus lunifer Westwood藏南深山14

ⅩⅩⅢ．Lucanus thibetanus furcifer Arrow藏深山云南亚种15

ⅩⅩⅣ．Lucanus thibetanus singularis Planet藏深山怒江亚种16

ⅩⅩⅤ．Lucanus thibetanus isaki Nagai藏深山独龙江亚种22

ⅩⅩⅥ．Lucanus thibetanus ssp藏深山片马未定亚种23

ⅩⅩⅦ．Lucanus langi Huang,He ＆ Shi郎氏深山23

ⅩⅩⅧ．Lucanus fryi Boileau弗瑞深山25

ⅩⅩⅨ．Lucanus atratus Hope阿深山25

ⅩⅩⅩ．Lucanus mearesii Hope金属深山26

ⅩⅩⅪ．Lucanus nyshiwini bretschneideri Schenk双齿深山墨脱亚种26

ⅩⅩⅫ．Eolucanus gracilis（Albers）短颚拟深山27

ⅩⅩⅩⅢ．Eolucanus lesnei（Planet）尖齿拟深山27

ⅩⅩⅩⅣ．Eolucanus mingyiae（Huang）圆齿拟深山27

ⅩⅩⅩⅤ．Eolucanus pani（Huang）潘氏拟深山30

ⅩⅩⅩⅥ．Eolucanus oberthuri（Planet）欧氏拟深山30

Further notes30

New descriptions30

ⅩⅩⅩⅦ．Lucanus choui Huang ＆ Chen sp．nov．周氏深山（新种）30

ⅩⅩⅩⅧ．Lucanus hewenjiae Huang ＆ Chen sp．nov．何氏深山（新种）33

Descriptions of some new taxa of Lucanidae from China35

A．Prismognathus zhangi Huang ＆ Chen sp．nov．张氏鬼锹（新种）35

B．Ceruchus yingqii Huang ＆ Chen sp．nov．浅纹铠锹（新种）36

C．Himaloaesalus Huang ＆ Chen gen．nov．喜马拉雅斑锹属（新属）39

Phylogeny and systematics of Lucaninae （sensu Holloway,1960 ＆ 1968）from China41

Introduction41

Abbreviations41

Choice of ingroup and outgroup taxa41

Source of data and initial names43

Morphology43

Characters coding and data sets45

Softwares46

Analyses and searching results47

Monophyla and higher taxa62

1） Genera62

2） Tribes64

3） Species groups in Lucanus65

4） Species groups in Neolucanus66

5） Species groups in Prosopocoilus,Dorcus and Falcicornis66

6） Clade of Sinodendron66

Systematics and annotated list of higher taxa of Chinese Lucanidae,with some extension to species from outside China,and with emphasis on the tribe Dorcini66

Unsorted genera from China66

Subfamily AESALINAE MacLeay,181967

Tribe Aesalini MacLeay,181967

Subfamily SYNDESINAE MacLeay,181967

Tribe Sinodendrini Mulsant,184267

Tribe Ceruchini Jacquelin du Val ＆ Fainnaire,185967

Subfamily LUCANINAE Latreille,180467

Tribe Platycerini Mulsant,184267

Tribe Noseolucanini Huang ＆ Chen,2013,new tribe67

Tribe Aegini Huang ＆ Chen,2013,new tribe68

Tribe Odontolabini Parry,187069

Tribe Figulini Burmeister,184769

Tribe Capreolucanini Huang ＆ Chen,2013,new tribe69

Tribe Cyclommatini Huang ＆ Chen,2013,new tribe70

Tribe Lucanini Latreille,180471

Tribe Dorcini Parry,186472

Genus Gnaphaloryx Burmeister,184772

Genus Kirchnerius Schenk,200972

Genus Metallactulus Ritsema,188573

Genus Epidorcus Seguy,195473

Genus Serrognathus Motschulsky,186174

Genus Hexarthrius Hope,184275

Genus Rhaetus Parry,186475

Genus Rhaetulus Westwood,187176

Genus Pseudorhaetus Planet,189976

Genus Prosopocoilus Hope ＆ Westwood,184576

Genus Miwanus Huang ＆ Chen,new genus77

Genus Falcicornis Planet,189478

Genus Sinodorcus Huang ＆ Chen,new genus78

Genus Aulacostethus Waterhouse,186979

Genus Dorcus MacLeay,181979

Discussion on some tribes from outside China80

Key to tribes of Lucaninae from China82

Appendix 1-List of characters used for phylogenetic analyses84

Appendix 2-Data matrix101

A review of Dorcini（sensu Huang ＆ Chen,2013）from China115

Introduction115

Key to genera of Dorcini males115

Key to genera of Dorcini females117

Genus Gnaphaloryx Burmeister,1847黑锈锹属119

1．Gnaphaloryx opacus Burmeister,1847黑锈锹119

Genus Kirchnerius Schenk,2009钳锹属121

Key to Chinese species of Kirchnerius males121

Key to Chinese species of Kirchnerius females122

Kirchnerius spencei group122

2．Kirchnerius spencei（Hope,1840）comb．nov．钳锹122

Kirchnerius boreli group125

3．Kirchnerius guangxii Schenk,2009 广西钳锹125

4．Kirchnerius yangi（Fukinuki,2004）comb．nov．杨氏钳锹127

5．Kirchnerius simianshanus（Huang ＆ Chen,2011）comb．nov．四面山钳锹128

6．Kirchnerius boreli（Boileau,1904）comb．nov．印度钳锹129

Genus Metallactulus Ritsema,1885姬扁锹属131

7．Metallactulus parvulus（Hope,1845）姬扁锹131

Genus Epidorcus Séguy,1954前锹属132

Key to Chinese species of Epidorcus males132

Key to Chinese species of Epidorcus females133

8．Epidorcus gracilis（Saunders,1854）狭长前锹134

9．Epidorcus piceipennis（Westwood,1855）拟狭长前锹137

10．Epidorcus denticulatus（Boileau,1901）,comb．nov．单基齿前锹139

11．Epidorcus rondoni（Bomans ＆ Lacroix,1970）,comb．nov．短基齿前锹144

12．Epidorcus tonkinensis（Pouillaude,1913）,comb．nov．并基齿前锹145

13．Epidorcus bidentatus（Bomans,1978）,comb．nov．双基齿前锹147

Genus Serrognathus Motschulsky,1861扁锹属147

Key to Chinese taxa of Serrognathus males147

Key to Chinese taxa of Serrognathus females148

14．Serrognathus titanus（Boisduval,1835）扁锹149

Serrognathus titanus platymelus（Saunders,1854）扁锹典型亚种149

Serrognathus titanus fafner（Kriesche,［1921］）扁锹越南亚种154

Serrognathus titanus typhoniformis（Nagel,1924）扁锹云贵亚种155

Serrognathus titanus sika（Kriesche,［1921］）扁锹台湾亚种156

Serrognathus titanus castanicolor Motschulsky,1861扁锹东北亚种157

Serrognathus titanus ssp．incert．1扁锹版纳亚种159

Serrognathus titanus ssp．incert．2扁锹墨脱亚种160

Species delimitation161

Diagnosis for species163

15．Serrognathus consentaneus（Albers,1886）尖腹扁锹（别名：细齿扁）165

16．Serrognathus kyanrauensis（Miwa,1934）深山扁锹168

Genus Hexarthrius Hope,1842六节锹属169

Key to Chinese taxa of Hexarthrius males169

Key to Chinese taxa of Hexarthrius females170

17．Hexarthrius aduncus Jordan,1894印度六节锹170

18．Hexarthrius vitalisi Didier,1925维氏六节锹176

Hexarthrius vitalisi vitalisi Didier,1925维氏六节锹指名亚种176

Hexarthrius vitalisi tsukamotoi Nagai,1998维氏六节锹桂北亚种177

Hexarthrius vitalisi ssp．incert．1维氏六节锹桂西亚种178

Hexarthrius vitalisi ssp．incert．2维氏六节锹高黎贡山亚种178

Species delimitation179

Subspecies179

19．Hexarthrius partyi Hope,1842橘背六节锹（别名：橘背叉角、帕瑞六节）179

20．Hexarthrius forsteri（Hope,1840）佛氏六节锹182

21．Hexarthrius melchioritis Séguy,1954缅甸六节锹183

Genus Rhaetus Parry,1864巨鹿锹属185

22．Rhaetus westwoodi（Parry,1862）巨鹿锹（别名：金刚鹿）185

Genus Rhaetulus Westwood,1871鹿角锹属187

23．Rhaetulus crenatus Westwood,1871鹿角锹187

Rhaetulus crenatus crenatus Westwood,1871鹿角锹指名亚种187

Rhaetulus cretatus rubrifemoratus Nagai,2000鹿角锹华南亚种189

Rhaetulus crenatus boileaui Didier,1925？鹿角锹版纳亚种190

Rhaetulus crenatus ssp．incert．鹿角锹中缅亚种191

Species delimitation191

Subspecies191

Genus Pseudorhaetus Planet,1899拟鹿锹属192

Key to Chinese taxa of Pseudorhaetus192

24．Pseudorhaetus sinicus（Boileau,1899）拟鹿锹（别名：漆黑鹿角）192

Pseudorhaetus sinicus sinicus（Boileau,1899）拟鹿锹指名亚种192

Pseudorhaetus sinicus concolor Benesh,1960拟鹿锹台湾亚种193

Species delimitation194

Diagnosis for species194

25．Pseudorhaetus oberthuri Planet,1899红背拟鹿锹194

Genus Prosopocoilus Hope,1845锯锹属195

Key to Chinese species of Prosopocoilus males195

Key to Chinese species of Prosopocoilus females197

Prosopocoilus buddha group199

26．Prosopocoilus approximatus（Parry,1864）伪布达锯锹199

27．Prosopocoilus thibeticus（Westwood,1855）滇东南锯锹203

28．Prosopocoilus porrectus Bomans,1978歧齿锯锹206

29．Prosopocoilus fulgens（Didier,1927）毛刷锯锹206

30．Prosopocoilus confucius（Hope,1842）孔夫子锯锹208

31．Prosopocoilus giraffa giraffa（Olivier,1789）长颈鹿锯锹指名亚种209

32．Prosopocoilus politus（Parry,1862）亮锯锹210

33．Prosopocoilus forficula（Thomson,1856）圆翅锯锹212

Prosopocoilus astacoides group214

34．Prosopocoilus astacoides（Hope,1840）两点锯锹214

Species delimitation and subspecies delimitation214

Prosopocoilus astacoides astacoides（Hope,1840）两点锯锹指名亚种217

Prosopocoilus astacoides castaneus（Hope,1845）两点锯锹樟木亚种219

Prosopocoilus astacoides fraternus（Hope,1845）两点锯锹滇南亚种221

Prosopocoilus astacoides kachinensis Bomans ＆ Miyashita,1997两点锯锹藏东南亚种223

Prosopocoilus astacoides blanchardi（Parry,1873）两点锯锹普通亚种224

Prosopocoilus astacoides dubernardi（Planet,1899）两点锯锹滇西北亚种226

Other subspecies227

Diagnosis for species227

Prosopocoilus impressus（Waterhouse,1869）-Valid species,not found in China229

Prosopocoilus laterinus（Didier,1928）-Not found in China230

35．Prosopocoilus reni Huang ＆ Chen,2011任氏锯锹231

36．Prosopocoilus yazakii Nagai,2005三色锯锹231

Prosopocoilus inclinatus group232

37．Prosopocoilus inclinatus inclinatus（Motschulsky,［1858］）东北锯锹232

38．Prosopocoilus motschulskyi（Waterhouse,1869）高砂锯锹234

39．Prosopocoilus oweni melli Kriesche,1922欧文锯锹中国亚种236

40．Prosopocoilus doris Kriesche,［1921］平齿锯锹238

41．Prosopocoilus suturalis（Olivier,1789）丫纹锯锹240

42．Prosopocoilus biplagiatus（Westwood,1855）黄纹锯锹241

43．Prosopocoilus spineus（Didier,1927）红背锯锹243

Prosopocoilus maclellandi group245

44．Prosopocoilus jenkinsi（Westwood 1848）毛胫四点锯锹245

45．Prosopocoilus maclellandi（Hope,1842）四点锯锹249

46．Prosopocoilus laterotarsus（Houlbert,1915）砂缘锯锹251

Genus Miwanus Huang ＆ Chen,2013 gen．nov．双钩锹属（新属）254

47．Miwanus formosanus（Miwa,1929）comb．nov．双钩锹254

Miwanus formosanus formosanus（Miwa,1929）双钩锹指名亚种（台湾亚种）254

Miwanus formosanus capricornus（Didier,1931）双钩锹越南亚种255

Miwanus formosanus kishidai（Fujita,2010）双钩锹广东亚种256

Miwanus formosanus ssp．incert．双钩锹华中未定名亚种256

Species delimitation257

Genus Falcicornis Planet,1894小刀锹属257

Key to Chinese species of Falcicornis males257

Key to Chinese species of Falcicornis females259

48．Falcicornis neolucanoides sp．nov．圆翅小刀锹（新种）261

Falcicornis elegantulus group262

49．Falcicornis pseudaxis（Didier,1926）comb．nov．南红小刀锹262

50．Falcicornis mellianus（Kriesche,［1921］）comb．nov．黄毛小刀锹264

51．Falcicornis seguyi（De Lisle,1955）comb．nov．叉齿小刀锹266

Falcicornis vernicatus group269

52．Falcicornis humilis（Arrow 1935）comb．nov．西藏小刀锹269

53．Falcicornis vernicatus（Arrow,1938）comb．nov．戟小刀锹276

54．Falcicornis itoi（Bomans,1993）comb．nov．越南小刀锹279

55．Falcicornis taibaishanensis（Schenk,2008）comb．nov．拟戟小刀锹279

56．Falcicornis heyangi Huang ＆ Chen sp．nov．何洋小刀锹（新种）280

Falcicornis bisignatus group281

57．Falcicornis tenuecostatus（Fairmaire,1888）comb．nov．皮氏小刀锹281

Falcicornis tenuecostatus tenuecostatus（Fairmaire,1888）皮氏小刀锹华北亚种281

Falcicornis tenuecostatus pieli（Didier ＆ Séguy,1953）stat．nov．皮氏小刀锹华南亚种283

Falcicornis tenuecostatus mochizukii（Miwa,1937）stat．nov．皮氏小刀锹台湾亚种（望月锹）286

Falcicornis tenuecostatus ssp．incert．皮氏小刀锹西康亚种286

Species delimitation and diagnosis for species287

58．Falcicornis moellenkampi（Nagel,1924）comb．nov．莫氏小刀锹290

Falcicornis moellenkampi moellenkampi（Nagel,1924）莫氏小刀锹指名亚种290

Falcicornis moellenkampi bomansi（Lacroix,1981）stat．nov．莫氏小刀锹滇西亚种291

Diagnosis for species293

Records294

59．Falcicornis ruficrus（De Lisle,1970）红河小刀锹294

60．Falcicornis bisignatus（Parry,1862）束胸小刀锹295

61．Falcicornis fulvonotatus（Parry,1862）斑胸小刀锹300

62．Falcicornis rufonotatus （Pouillaude,1913）滇越小刀锹302

63．Falcicornis songianus（Didier ＆ Séguy,1953）华南小刀锹304

64．Falcicornis himalayae（Schenk 2009）宽胸小刀锹307

Genus Sinodorcus Huang ＆ Chen,2013 gen．nov．多刺刀锹属（新属）308

65．Sinodorcus sawaii（Tsukawaki,1999）comb．nov．多刺刀锹308

Genus Aulacostethus Waterhouse,1869阿锹属309

Key to species of Aulacostethus309

66．Aulacostethus archeri Waterhouse,1869印度阿锹310

67．Aulacostethus ruditemporalis（Houlbert,1914）四川阿锹311

68．Aulacostethus doani Baba,2000越南阿锹312

69．Aulacostethus tianmuxing Huang ＆ Chen sp．nov．华东阿锹（新种）313

Genus Dorcus MacLeay,1819刀锹属、大锹属314

Key to Chinese species of Dorcus males315

Key to Chinese species of Dorcus females326

Dorcus antaeus group333

70．Dorcus schenklingi（M？llenkamp,1913）长角大锹333

71．Dorcus antaeus Hope,1842安达大锹（别名：安达佑实大锹）335

72．Dorcus vicinus Saunders,1854华东大锹（别名：天章大锹）339

73．Dorcus rectus（Motschulsky,［1858］）单齿刀锹（别名：日本小锹、直牙锹）341

Dorcus montivagus group344

74．Dorcus davidis（Fairmaire,1887）大卫大锹344

Dorcus rubrofemoratus group348

75．Dorcusyamadai（MIWA,1937）台湾刀锹（别名：刀锹）348

76．Dorcus rubrofemoratus（van Vollenhoven,1865）红腿刀锹351

Dorcus rubrofemoratus rubrofemoratus（van Vollenhoven,1865）红腿刀锹指名亚种351

Dorcus rubrofemoratus chenpengi（Li,1992）红腿刀锹北部亚种352

Dorcus rubrofemoratus miyamai Nagai ＆ Okazaki,2005红腿刀锹西部亚种353

Species delimitation and diagnosis for species354

77．Dorcus haitschunus（Didier ＆ Séguy,1952）长齿刀锹354

78．Dorcus arrowi（Boileau,1911）红背刀锹356

Dorcus arrowi arrowi（Boileau,1911）红背刀锹指名亚种（滇西亚种）356

Dorcus arrowi magdaleinae（Lacroix,1972）stat．nov．红背刀锹滇南亚种358

Dorcus arrowi ssp．incert．红背刀锹滇东南亚种360

Diagnosis for species362

79．Dorcus derelictus Parry,1862短颚刀锹363

Dorcus nepalensis group364

80．Dorcus sinensis（Boileau,1899）中华刀锹364

81．Dorcus semenowi Jakowlew,1900谢苗诺夫刀锹371

82．Dorcus kikunoae Hosoguchi,2004错那刀锹377

83．Dorcus wui Huang ＆ Chen sp．nov．吴氏刀锹（新种）378

84．Dorcus ratiocinativus Westwood,1871弯颚刀锹380

85．Dorcus wardi Arrow,1943沃德刀锹381

86．Dorcus macleayii（Hope,1845）麦氏刀锹383

87．Dorcus branaungi Nagai,2000布氏刀锹（别名：布朗刀）386

88．Dorcus sp．incert．1二郎山刀锹387

89．Dorcus sp．incert．2巨刀锹388

90．Dorcus nepalensis（Hope,1831）尼泊尔刀锹389

91．Dorcus donckieri（Boileau,1898）东氏刀锹392

Dorcus curvidens group393

92．Dorcus curvidens（Hope,1840）弯角大锹393

93．Dorcus hopei（Saunders,1854）中华大锹（别名：中国大锹）397

Dorcus hopei hopei（Saunders,1854）中华大锹指名亚种397

Dorcus hopei formosanus Miwa,1929 stat．nov．中华大锹台湾亚种400

Diagnosis for species401

Subspecies402

94．Dorcus yaksha Gravely,1915细角大锹402

Dorcus yaksha yaksha Gravely,1915细角大锹指名亚种402

Dorcus yaksha gracilicornis Benesh,1950 stat．nov．细角大锹东部亚种404

Dorcus katsurai group407

95．Dorcus katsurai Ikeda,2000扩胫大锹408

96．Dorcus sukkiti Fukinuki,2004缅甸扩胫大锹411

97．Dorcus sp．incert．3肘子大锹411

98．Dorcus myinti Nagai ＆ Maeda,2009敏氏刀锹411

Dorcus velutinus group413

99．Dorcus velutinus Thomson,1862锈刀锹413

100．Dorcus tenuihirsutus Kim ＆ Kim,2010北方锈刀锹418

101．Dorcus ursulus Arrow 1938扩胸锈刀锹419

102．Dorcus taiwanicus Nakane ＆ Makino,1985台湾锈刀锹420

103．Docus carinulatus Nagel,1941直颚锈刀锹422

104．Dorcus ursulae（Schenk,1996）平齿刀锹424

105．Dorcus tanakai Nagai,2002田中氏刀锹426

Dorcus elegans group427

106．Dorcus elegans（Parry,1862）樟木怪刀锹427

107．Docus motuoensis Huang ＆ Chen sp．nov．墨脱怪刀锹（新种）429

108．Dorcus chucheni Huang ＆ Chen sp．nov．初氏怪刀锹（新种）433

109．Dorcus wemckeni（Schenk,2008）短颚怪刀锹434

Dorcus striatipennis group（sensu stricto）435

110．Dorcus striatipennis（Motschulsky,1861）双齿刀锹435

Dorcus striatipennis yushiroi Sakaino,1997双齿刀锹台湾亚种435

Dorcus striatipennis davidi（Séguy,1954）stat．nov．双齿刀锹大陆亚种436

Delimitation of species439

Subspecies441

111．Dorcus lvbu Huang ＆ Chen sp．nov．吕布刀锹（新种）442

112．Dorcus bashanus Huang ＆ Chen sp．nov．巴山刀锹（新种）445

113．Dorcus gongshanus Huang ＆ Chen sp．nov．贡山刀锹（新种）446

114．Dorcus intricatus（Lacroix,1981）雅安刀锹448

115．Dorcus linwenhsini Huang ＆ Chen sp．nov．林氏刀锹（新种）450

Dorcus lhoba group452

116．Dorcus lhoba Huang ＆ Chen sp．nov．珞巴大锹（新种）452

117．Dorcus menba Huang ＆ Chen sp．nov．门巴大锹（新种）455

Dorcus zhangmuensis group456

118．Dorcus zhangmuensis Huang ＆ Chen sp．nov．樟木大锹（新种）456

119．Dorcus nosei Nagai,2000察隅大锹457

120．Dorcus cuonaensis Huang ＆ Chen sp．nov．错那大锹（新种）460

Dorcus reichei group462

121．Dorcus mencius（Kriesche,1935）孟子大锹463

122．Dorcus liyingbingi Huang ＆ Chen sp．nov．李氏大锹（新种）467

123．Dorcus reichei Hope,1842瑞奇大锹469

124．Dorcus cervulus（Boileau,1901）拟瑞奇大锹485

125．Dorcus hirticornis（Jakowlew,［1897］）毛角大锹487

Dorcus hirticornis hirticornis（Jakowlew,［1897］）毛角大锹指名亚种487

Dorcus hirticornis clypeatus Benesh,1950 stat．nov．毛角大锹台湾亚种488

Dorcus hirticornis ssp．毛角大锹云南亚种491

Species delimitation and diagnosis for species492

Dorcus hansteini（Albers,1889）----not found in China507

126．Dorcus kyawi Nagai ＆ Maeda,2009久氏大锹509

Dorcus tityus group511

127．Dorcus lineatopunctatus（Hope 1831）尼泊尔大锹511

Dorcus lineatopunctatus gaoligongshanus subsp．nov．尼泊尔大锹东部亚种（新亚种）520

Basic information of males521

128．Dorcus tityus Hope,1842提提大锹（别名：提提乌斯大锹）527

129．Dorcus daedalion（Didier ＆ Séguy,1953）华南大锹531

Dorcus submolais（Hoper,1845）----not found in China536

Dorcus fuliginosus（Didier,1928）----not found in China538

Dorcus hyperion group539

130．Dorcus laevidorsis Fairmaire,1888滇越大锹539

131．Dorcus hyperion Boileau,1899海波力昂大锹544

132．Dorcus hansi Schenk 2008基齿大锹（别名：三牙大锹）548

133．Dorcus miwai Benesh,1936平头大锹550

Unreliable records of Dorcini species for Chinese fauna in references551

References cited in all chapters of this book553

Plates569

Index to species-group names,genus-group names and family-group names709